Julien, Maurice and Pancho regroup aboard a submarine and ponder their next move, while a captive Mort plots to take down the tyrannical new king.
Clover hopes Sage's guru can help with a plan to oust Koto, who's now made Mort his jester. Julien and crew meet an enchanting tribe of lemur sirens.
Mort leads an effort to unseat Koto. Julien, Maurice and Pancho risk being fed to a tentacled monster. Sage's guru is not quite what Clover expected.
As Julien and company head back to Madagascar, their submarine runs into trouble. Mort puts on a play for Koto to cover up the captives' escape.
Julien and the boys team up with sharks and try to form an army. Jarsh-Jarsh's map leads Clover and Sage into danger. Mort's escape plan hits a snag.
A band of disgruntled Russian space chimps may be Julien's best hope for fighting Koto. Mort finds Timo locked in his pod and begs him for help.
Clover rethinks her decision to join Pam's cult. Mort uses Timo's radio to contact Julien and company, who are now in trouble with the chimps.
Donning disguises, Julien and friends try to hitch a ride on a Foosa ship. Jarsh-Jarsh presents Clover with a confusing new challenge.
Julien and Maurice face a harrowing fight at gladiator school. Mort travels to another dimension. Sage convinces Clover there's still good in Koto.
A colony of bell-shaped creatures hails Maurice as their Chosen One. Stuck in limbo, Julien stands trial for his life. Mort battles Morticus Khan.
Julien and Karl set out to stop Koto's wedding to the Crocodile Princess. Clover deals with Koto her own way. Mort fights to bring back "Smart Mort."
As Koto prepares a sacrifice to kick off his wedding, Clover and Sage unleash their new weapon. Mort, Maurice and Pancho arrive with reinforcements.
Nearly vanquished by Koto's troops, Julien rallies Mort's army, the bell snails, the space chimps, Clover and other loyal friends for one last hurrah.